The largest organisms found on Earth can be determined according to various aspects of an organism's size, such as: mass, volume, area, length, height, or even genome size. Some organisms group together to form a superorganism (such as ants or bees), but such are not classed as single large organisms. The Great Barrier Reef is the world's largest structure composed of living entities, stretching 2,000Â km (1,200Â mi), but contains many organisms of many types of species.
This article lists the largest species for various types of organisms, and mostly considers extant species. The organism sizes listed are frequently considered "outsized" and are not in the normal size range for the respective group.
If considered singular entities, the largest organisms are clonal colonies which can spread over large areas. Pando, a clonal colony of the aspen tree, is widely considered to be the largest such organism by mass. Even if such colonies are excluded, trees retain their dominance of this listing, with the giant sequoia being the most massive tree. In 2006 a huge clonal colony of Posidonia oceanica was discovered south of the island of Ibiza. At 8 kilometres (5.0Â mi) across, and estimated at around 100,000 years old, it may be one of the largest and oldest clonal colonies on Earth.
Among animals, the largest species are all marine mammals, specifically whales. The blue whale is believed to be the largest animal to have ever lived. The largest land animal classification is also dominated by mammals, with the African bush elephant being the most massive of these.
Plants
The largest single-stem tree by wood volume and mass is the giant sequoia (Sequoiadendron giganteum), native to Sierra Nevada and California; it grows to an average height of 70â"85Â m (230â"280Â ft) and 5â"7Â m (16â"23Â ft) in diameter.
Multiple-stem trees such as banyan can be enormous. Thimmamma Marrimanu in India spreads over 1.0Â ha (2.5 acres).
The largest organism in the world, according to mass, is the aspen tree whose colonies of clones can grow up to five miles long.
Another form of flowering plant that rivals Pando as the largest organism on earth in breadth, if not mass, is the giant marine plant, Posidonia oceanica, discovered in the Mediterranean near the Balearic Islands, Spain. Its length is about 8Â km (5.0Â mi). It may also be the oldest living organism in the world, with an estimated age of 100,000 years.
Animals
A member of the order Cetacea, the blue whale (Balaenoptera musculus), is thought to be the largest animal ever to have lived. The maximum recorded weight was 190 metric tonnes for a specimen measuring 30 metres (98Â ft), whereas longer ones, up to 33.4 metres (110Â ft), have been recorded but not weighed.
The African bush elephant (Loxodonta africana), of the order Proboscidea, is the largest living land animal. A native of various open habitats in sub-Saharan Africa, this elephant is born commonly weighing about 100 kilograms (220Â lb). The largest elephant ever recorded was shot in Angola in 1974. It was a male measuring 10.7 metres (35Â ft) from trunk to tail and 4.17 metres (13.7Â ft) lying on its side in a projected line from the highest point of the shoulder to the base of the forefoot, indicating a standing shoulder height of 3.96 metres (13.0Â ft).
- Table of heaviest living animals
The heaviest living animals are all cetaceans, and thus also the largest living mammals. Since no scale can accommodate the whole body of a large whale, most whales have been weighed by parts.
- Table of heaviest terrestrial animals
The following is a list of the heaviest wild land animals, which are all mammals. The African elephant is now listed as two species, the African bush elephant and the African forest elephant, as they are now generally considered to be two separate species.
Tunicates (Tunicata)
The largest tunicates are Synoicum pulmonaria, found at depths of 20 and 40 metres (66 and 131Â ft), and are up to 14 centimetres (6Â in) in diameter. It is also present in the northwestern Atlantic Ocean, around the coasts of Greenland and Newfoundland, but is less common here than in the east, and occurs only at depths between 10 and 13 metres (33 and 43Â ft).
- Entergonas (Enterogona)
- The largest entergonas Synoicum pulmonaria it is usually found at depths between about 20 and 40 metres (66 and 131Â ft) and can grow to over a metre (yard) in length. It is also present in the northwestern Atlantic Ocean, around the coasts of Greenland and Newfoundland, but is less common here than in the east, and occurs only at depths between 10 and 13 metres (33 and 43Â ft).
- Pleurogonas (Pleurogona)
- The largest pleurogonas: Pyura pachydermatina . In colour it is off-white or a garish shade of reddish-purple. The stalk is two thirds to three quarters the length of the whole animal which helps distinguish it from certain invasive tunicates not native to New Zealand such as Styela clava and Pyura stolonifera. It is one of the largest species of tunicates and can grow to over a metre (yard) in length.
- Aspiraculates (Aspiraculata)
- The largest aspiraculates: Oligotrema large and surrounded by six large lobes; the cloacal syphon is small. They live exclusively in deep water and range in size from less than one inch (2 cm) to 2.4 inches (6 cm).
Thaliacea (Thaliacea)
- The largest thaliacean: Pyrosoma atlanticum is cylindrical and can grow up to 60 cm (2 ft) long and 4â"6 cm wide. The constituent zooids form a rigid tube, which may be pale pink, yellowish, or bluish. One end of the tube is narrower and is closed, while the other is open and has a strong diaphragm. The outer surface or test is gelatinised and dimpled with backward-pointing, blunt processes. The individual zooids are up to 8.5 mm (0.3 in) long and have a broad, rounded branchial sac with gill slits. Along the side of the branchial sac runs the endostyle, which produces mucus filters. Water is moved through the gill slits into the centre of the cylinder by cilia pulsating rhythmically. Plankton and other food particles are caught in mucus filters in the processes as the colony is propelled through the water. P. atlanticum is bioluminescent and can generate a brilliant blue-green light when stimulated.
- Doliolida (Doliolida)
- The largest doliolida: Doliolida The doliolid body is small, typically 1â"2Â cm long, and barrel-shaped; it features two wide siphons, one at the front and the other at the back end, and eight or nine circular muscle strands reminiscent of barrel bands. Like all tunicates, they are filter feeders. They are free-floating; the same forced flow of water through their bodies with which they gather plankton is used for propulsion - not unlike a tiny ramjet engine. Doliolids are capable of quick movement. They have a complicated lifecycle consisting of sexual and asexual generations. They are nearly exclusively tropical animals, although a few species can be found as far to the north as northern California.
- Salps (Salpida)
- The largest salps: Cyclosalpa bakeri15cm (6ins) long. There are openings at the anterior and posterior ends of the cylinder which can be opened or closed as needed. The bodies have seven transverse bands of muscle interspersed by white, translucent patches. A stolon grows from near the endostyle (an elongated glandular structure producing mucus for trapping food particles). The stolon is a ribbon-like organ on which a batch of aggregate forms of the animal are produced by budding. The aggregate is the second, colonial form of the salp and is also gelatinous, transparent and flabby. It takes the shape of a radial whorl of individuals up to about 20cm (4in) in diameter. It is formed of approximately 12 zooids linked side by side in a shape that resembles a crown. are largest thetyses: Thetys vagina Individuals can reach up to 30Â cm (12Â in) long.
- Larvaceans (Larvacea)
- The largest larvaceans: Appendicularia 1Â cm (0.39Â in) in body length (excluding the tail).
Cephalochordate (Leptocardii)
The largest lancelets: European lancelet (Branchiostoma lanceolatum) "primitive fish". It can grow up to 6Â cm (2.5Â in) long.
Vertebrates
Mammals (Mammalia)
The blue whale is the largest mammal.
The largest land mammal extant today is the African bush elephant. The largest extinct species known was once considered to be Paraceratherium orgosensis, a rhinoceros relative thought to have stood up to 5.5Â m (18Â ft) tall, measured over 9Â m (30Â ft) long and may have weighed up to 20 tonnes. More recent estimates suggest that Paraceratherium was surpassed by the proboscidean Palaeoloxodon namadicus at about 22 tonnes.
Stem-mammals (Synapsida)
The Permian era Cotylorhynchus, from what is now the southern United States, probably was the largest of all synapsids (most of which became extinct 250 million years ago), at 6Â m (20Â ft) and 2 tonnes. The largest carnivorous synapsid was Anteosaurus from what is now South Africa during Middle Permian era. Anteosaurus was 5â"6Â m (16â"20Â ft) long, and weighed about 500â"600Â kg (1,100â"1,300Â lb).
- Pelycosauria
- The largest pelycosaur was the pre-mentioned Cotylorhynchus, and the largest predatory pelycosaurus was Dimetrodon grandis from what is now North America, with a length of 3.1Â m (10Â ft) and weight of 250Â kg (550Â lb).
- Therapsida
- Moschops was the largest non-mammalian therapsid, with a weight of 700 to 1,000Â kg (1,500 to 2,200Â lb), and a length of about 5Â m (16Â ft). The largest carnivorous therapsid was the aforementioned Anteosaurus.
Reptiles (Reptilia)
The largest living reptile, a representative of the order Crocodilia, is the saltwater crocodile (Crocodylus porosus) of Southern Asia and Australia, with adult males being typically 3.9â"5.5Â m (13â"18Â ft) long. The largest confirmed saltwater crocodile on record was 6.32Â m (20.7Â ft) long, and weighed about 1,360Â kg (3,000Â lb). Unconfirmed reports of much larger crocodiles exist, but examinations of incomplete remains have never suggested a length greater than 7Â m (23Â ft). Also, a living specimen estimated at 7Â m (23Â ft) and 2,000Â kg (4,400Â lb) has been accepted by the Guinness Book of World Records. However, due to the difficulty of trapping and measuring a very large living crocodile, the accuracy of these dimensions has yet to be verified. A specimen named Lolong caught alive in the Philippines in 2011 (died February 2013) was found to have measured 6.17Â m (20.2Â ft) in length.
The Komodo dragon (Varanus komodoensis), also known as the "Komodo monitor", is a large species of lizard found in the Indonesian islands of Komodo, Rinca, Flores, Gili Motang, and Padar. A member of the monitor lizard family (Varanidae), it is the largest living species of lizard, growing to a maximum length of 3 metres (9.8 feet) in rare cases and weighing up to approximately 70 kilograms (150 pounds).
- Table of heaviest living reptiles
The following is a list of the heaviest living reptile species ranked by average weight, which is dominated by the crocodilians. Unlike mammals, birds or fish, the mass of large reptiles is frequently poorly documented and many are subject to conjecture and estimation.
Dinosaurs (Dinosauria)
- Now extinct, except for birds, which are theropods.
- Sauropods (Sauropoda)
- The largest dinosaurs, and the largest animals to ever live on land, were the plant-eating, long-necked Sauropoda. The tallest and heaviest sauropod known from a complete skeleton is a specimen of an immature Giraffatitan discovered in Tanzania between 1907 and 1912, now mounted in the Museum für Naturkunde of Berlin. It is 12 m (39 ft) tall and weighed 23.3â"39.5 tonnes. The longest is a 25 m (82 ft) long specimen of Diplodocus discovered in Wyoming, and mounted in Pittsburgh's Carnegie Natural History Museum in 1907. A Patagotitan specimen found in Argentina in 2014 is estimated to have been 40 m (130 ft) long and 20 m (66 ft) tall, with a weight of 77 tonnes.
- There were larger sauropods, but they are known only from a few bones. The current record-holders include Argentinosaurus, which may have weighed 73 tonnes; Supersaurus which might have reached 34Â m (112Â ft) in length and Sauroposeidon which might have been 18Â m (59Â ft) tall. Two other such sauropods include Bruhathkayosaurus and Amphicoelias fragillimus. Both are known only from fragments. Bruhathkayosaurus might have been between 40â"44Â m (131â"144Â ft) in length and 175â"220 tonnes in weight according to some estimates. A. fragillimus might have been approximately 58Â m long and 122.4Â metric tons in weight.
- Theropods (Theropoda)
- The largest theropod known from a nearly complete skeleton is the biggest and most complete Tyrannosaurus rex specimen, nicknamed "Sue", which was discovered in South Dakota in 1990 and now mounted in the Field Museum of Chicago at a total length of 12.3Â m (40Â ft). Body mass estimates have reached over 9500Â kg, though other figures, such as Hartmanâs 2013 estimate of 8400Â kg, have been lower.
- Another giant theropod is the semi-aquatic Spinosaurus aegyptiacus from the mid-Cretaceous of North Africa. Size estimates have been fluctuating far more over the years, with length estimates ranging from 12.6 to 18 m and mass estimates from 7 to 20.9 t. Recent findings favour a length exceeding 15 m and a body mass of 7.5 tons.
- Other contenders known from partial skeletons include Giganotosaurus carolinii (est. 12.2â"13.2 m and 6-13.8 tonnes) and Carcharodontosaurus saharicus (est. 12-13.3 m and 6.2-15.1 tonnes).
- The largest extant theropod is the Ostrich (see birds, below).
- Armored dinosaurs (Thyreophora)
- The largest thyreophorans were Ankylosaurus and Stegosaurus, from the Late Cretaceous and Late Jurassic periods (respectively) of what is now North America, both measuring up to 9Â m (30Â ft) in length and estimated to weigh up to 6 tonnes.
- Ornithopods (Ornithopoda)
- The largest ornithopods, were the hadrosaurids Shantungosaurus, a late Cretaceous dinosaur found in the Shandong Peninsula of China, and Magnapaulia from the late Cretaceous of North America. Both species are known from fragmentary remains but are estimated to have reached over 15Â m (49Â ft) in length and were likely the heaviest non-sauropod dinosaurs, estimated at over 23Â tonnes.
- Ceratopsians (Ceratopsia)
- The largest ceratopsians were Triceratops and its ancestor Eotriceratops from the late Cretaceous of North America. Both estimated to have reached about 9Â m (30Â ft) in length and weighed 12Â tonnes.
Birds (Aves)
The largest living bird, a member of the Struthioniformes, is the common ostrich (Struthio camelus), from the plains of Africa and Arabia. A large male ostrich can reach a height of 2.8Â m (9.2Â ft) and weigh over 156Â kg (344Â lb). A mass of 200Â kg (440Â lb) has been cited for the common ostrich but no wild ostriches of this massive weight have been verified. Eggs laid by the ostrich can weigh 1.4Â kg (3.1Â lb) and are the largest eggs in the world today.
The largest bird in the fossil record may be the extinct elephant birds (Aepyornis) of Madagascar, which were related to the ostrich. They exceeded 3Â m (9.8Â ft) in height and 500Â kg (1,100Â lb). The last of the elephant birds became extinct about 300 years ago. Of almost exactly the same upper proportions as the largest elephant birds was Dromornis stirtoni of Australia, part of a 26,000-year-old group called mihirungs of the family Dromornithidae. The largest carnivorous bird was Brontornis, an extinct flightless bird from South America which reached a weight of 350 to 400Â kg (770 to 880Â lb) and a height of about 2.8Â m (9Â ft 2Â in). The tallest carnivorous bird was Kelenken, which could reach 3 to 3.2 meters in height and 220 to 250 kilograms. The tallest bird ever was the giant moa (Dinornis maximus), part of the moa family of New Zealand that went extinct around 1500 AD. This particular species of moa stood up to 3.7Â m (12Â ft) tall, but weighed about half as much as a large elephant bird or mihirung due to its comparatively slender frame.
The heaviest bird ever capable of flight was Argentavis magnificens, the largest member of the now extinct family Teratornithidae, found in Miocene-aged fossil beds of Argentina, with a wingspan up to 5.5Â m (18Â ft), a length of up to 1.25Â m (4.1Â ft), a height on the ground of up to 1.75Â m (5.7Â ft) and a body weight of at least 71Â kg (157Â lb). Pelagornis sandersi is thought to have had an even larger wingspan of about 6.1â"7.4Â m (20â"24Â ft), but is only about 22â"40Â kg (49â"88Â lb), half the mass of the former.
- Table of heaviest living birds
The following is a list of the heaviest living bird species ranked by average weight. These species are almost all flightless, which allows for these particular birds to have denser bones and heavier bodies. Flightless birds comprise less than 2% of all living bird species.
Amphibians (Amphibia)
The largest living amphibian is the Chinese giant salamander (Andrias davidianus). The maximum size of this nearly human-sized river-dweller is 64Â kg (141Â lb) and almost 1.83Â m (6.0Â ft). Before amniotes became the dominant tetrapods, several giant amphibian proto-tetrapods existed and were certainly the dominant animals in their ecosystems. The largest known was the crocodile-like Prionosuchus, which reached a length of 9Â m (30Â ft).
- Frogs (Anura)
- The largest member of the largest order of amphibians is the African Goliath frog (Conraua goliath). The maximum size this species is verified to attain is a weight of 3.8Â kg (8.4Â lb) and a snout-to-vent length of 39Â cm (15Â in). The largest of the toads, the cane toad (Rhinella marina), is also the second largest member of the frog order. This infamous, often invasive species can grow to maximum mass of 2.65Â kg (5.8Â lb) and measure a maximum of 33Â cm (13Â in) from snout-to-vent. Rivaling the previous two species, the African bullfrog (Pyxicephalus adspersus) can range up to a weight of 2Â kg (4.4Â lb) and 25.5Â cm (10.0Â in) from snout to vent. Another large frog is the largest frog in North America, the American bullfrog, which can reach weights of up to 0.8Â kg (1.8Â lb) and snout-to-vent-length (SVL) of 20Â cm (7.9Â in). However, the toad Beelzebufo ampinga, found in fossil from the Cretaceous era in what is now Madagascar, could grow to be 41Â cm (16Â in) long and weigh up to 4.5Â kg (9.9Â lb), making it the largest frog ever known. The largest tree frog is the Australasian white-lipped tree frog (Litoria infrafrenata), the females of which can reach a length of 14Â cm (5.5Â in) from snout to vent and can weigh up to 115Â g (4.1Â oz). The family Leptodactylidae, one of the most diverse anuran families, also has some very large members. The largest is the Surinam horned frog (Ceratophrys cornuta), which can reach 20Â cm (7.9Â in) in length from snout to vent and weigh up to 0.48Â kg (1.1Â lb). While not quite as large as Ceratophrys cornuta, Leptodactylus pentadactylus is often heavier; it can reach 18.5Â cm (7.3Â in) long and weigh 0.60 kilograms (1.3 pounds). The largest dendrobatid is the Colombian golden poison frog (Phyllobates terribilis), which can attain a length of 6Â cm (2.4Â in) and nearly 28.3Â g (1.00Â oz). Most frogs are classified under the suborder Neobatrachia, although nearly 200 species are part of the Mesobatrachia suborder, or ancient frogs. The largest of these are the little-known Brachytarsophrys or Karin Hills frogs, of South Asia, which can grow to a maximum snout-to-vent length of 17Â cm (6.7Â in) and a maximum weight of 0.54Â kg (1.2Â lb).
- Caecilians (Gymnophiona)
- The largest of the worm-like caecilians is the Colombian Thompson's caecilian (Caecilia thompsoni), which reaches a length of 1.5Â m (4.9Â ft), a width of about 4.6Â cm (1.8Â in) and can weigh up to about 1Â kg (2.2Â lb).
- Salamanders (Urodela)
- Besides the previously mentioned Chinese giant salamander, the closely related Japanese giant salamander (Andrias japonicus) is also sometimes cited as the largest living amphibian, but salamanders of a greater size than 1.53Â m (5.0Â ft) and 36Â kg (79Â lb) have never been verified for this species. Another giant of the amphibian world is the North American Hellbender (Cryptobranchus alleganiensis), which can measure up to 0.76Â m (2.5Â ft). The largest of the newts is the Iberian ribbed newt (Pleurodeles waltl), which can grow up to 30Â cm (12Â in) in length.
Fish
Invertebrates
Sponges (Porifera)
The largest known species of sea sponge is the giant barrel sponge, Xestospongia muta. These massively built sponges can reach 2.4Â m (8Â ft) in height and can be of about the same number of feet across at the thickest part of the "body". Some of these creatures have been estimated to be over 2,400 years of age.
- Calcareous sponges (Calcarea)
- The largest known of these small, inconspicuous sponges is probably the species Pericharax heteroraphis, attaining a height of 30Â cm (0.98Â ft). Most calcareous sponges do not exceed 10Â cm (3.9Â in) tall.
- Hexactinellid sponges (Hexactinellida)
- A relatively common species, Rhabdocalyptus dawsoni, can reach a height of 1Â m (3.3Â ft) once they are of a very old age. This is the maximum size recorded for a hexactinellid sponge.
Cnidarians (Cnidaria)
The lion's mane jellyfish (Cyanea capillata) is the largest cnidaria species, of the class Scyphozoa. The largest known specimen of this giant, found washed up on the shore of Massachusetts Bay in 1870, had a bell diameter of 2.5Â m (8.2Â ft), a weight of 150Â kg (330Â lb). The tentacles of this specimens were as long as 37Â m (121Â ft) and were projected to have a tentacular spread of about 75Â m (246Â ft) making it one of the longest extant animals.
- Corals and sea-anemones (Anthozoa)
- The largest individual species are the sea-anemones of the genus Discoma, which can attain a mouth disc diameter of 60Â cm (2.0Â ft). Longer, but much less massive overall, are the anemones of the genus Ceriantharia, at up to 2Â m (6.6Â ft) tall. Communities of coral can be truly massive, a single colony of the Porites genus can be over 10Â m (33Â ft), but the actual individual organisms are quite small.
- Lion's mane jellyfish
- The largest known species of jellyfish is the "lion's mane jellyfish". The largest of its kind was found in 1870 in Massachusetts Bay with a recorded bell size of 2.3 metres (7Â feet 7Â inches) and a tentacle length of 37 metres (121 feet).
- Hydrozoans (Hydrozoa)
- The colonial siphonophore Praya dubia can attain lengths of 40â"50Â m (130â"160Â ft). The Portuguese man o' war's (Physalia physalis) tentacles can attain a length of up to 50Â m (160Â ft).
Flatworms (Platyhelminthes)
The largest terrestrial species of this phylum of mostly parasites, commonly called flatworms, is the greenhouse planarian (Bipalium kewense). This planarian can reach a length of 60Â cm (2.0Â ft) and is quite massive for a flatworm.
- Monogenean flatworms (Monogenea)
- The largest known members of this group of very small parasites are among the genus of capsalids, Listrocephalos, reaching a length of 2Â cm (0.79Â in).
- Flukes (Trematoda)
- The largest known species of fluke is Fasciolopsis buski, which most often attacks humans and livestock. One of these flukes can be up to 7.5Â cm (3.0Â in) long and 2Â cm (0.79Â in) thick.
- Tapeworms (Cestoda)
- The largest known species of tapeworm is the whale tapeworm, Polygonoporus giganticus, which can grow to over 30Â m (98Â ft).
Roundworms (Nematoda)
The largest roundworm, Placentonema gigantissima, is a parasite found in the placentas of sperm whales which can reach up to 9Â m (30Â ft) in length.
Segmented worms (Annelida)
The largest of the segmented worms (commonly called earthworms) is the African giant earthworm (Microchaetus rappi). Although it averages about 1.36Â m (4.5Â ft) in length, this huge worm can reach a length of as much as 6.7Â m (22Â ft) and can weigh over 1.5Â kg (3.3Â lb). Only the giant Gippsland earthworm, Megascolides australis, and a few giant polychaetes reach nearly comparable sizes, reaching 4 and 3.6Â m (13 and 12Â ft), respectively.
Echinoderms (Echinodermata)
The largest species of echinoderm in terms of bulk is probably the starfish species Thromidia gigas, of the class Asteroidea, which reaches a weight of over 6Â kg (13Â lb), but it might be beaten by some giant sea cucumbers such as Thelenota anax. However, at a maximum span of 63Â cm (25Â in), Thromidia gigas is quite a bit shorter than some other echinoderms. The longest echinoderm known is the conspicuous sea cucumber Synapta maculata, with a slender body that can extend up to 3Â m (9.8Â ft). In comparison, the biggest sea star is the brisingid sea star Midgardia xandaros, reaching a span of 1.4Â m (4.6Â ft), despite being quite slender. Evasterias echinosoma is another giant echinoderm and can measure up to 1Â m (3.3Â ft) across and weigh 5.1Â kg (11Â lb).
- Crinoids (Crinoidea)
- The largest species of crinoid is the unstalked feather-star Heliometra glacialis, reaching a total width of 78Â cm (31Â in) and an individual arm length of 35Â cm (14Â in). A width of 91.4Â cm (36.0Â in) was claimed for one unstalked feather-star but is not confirmed. The genus Metacrinus has a stalk span of 61Â cm (24Â in) but, due to its bulk and multiple arms, it is heavier than Heliometra. In the past, crinoids grew much larger, and stalk lengths up to 40Â m (130Â ft) have been found in the fossil record.
- Sea urchins and allies (Echinoidea)
- The largest sea urchin is the species Sperosoma giganteum from the deep northwest Pacific Ocean, which can reach a shell width of about 30Â cm (12Â in). Another deep sea species Hygrosoma hoplacantha is only slightly smaller. The largest species found along the North America coast is the Pacific red sea urchin (Mesocentrotus franciscanus) where the shell can reach 19Â cm (7.5Â in). If the spines enter into count, the biggest species may be a Diadematidae like Diadema setosum, with a test up to 10Â cm (3.9Â in) only, but its spines can reach up to 30Â cm (12Â in) in length.
- Sea cucumbers (Holothuroidea)
- The bulkiest species of sea cucumber are Stichopus variegatus and Thelenota anax, weighing several pounds, being about 21Â cm (8.3Â in) in diameter, and reaching a length of 1Â m (3.3Â ft) when fully extended. Synapta maculata can reach an extended length of 3Â m (9.8Â ft), but is extremely slender (3-5cm) and weigh much less than Stichopodids.
- Brittle stars (Ophiuroidea)
- The largest known specimen of brittle star is the basket star Astrotoma agassizii. This species can grow to have a span of 1Â m (3.3Â ft). Sometimes, Gorgonocephalus stimpsoni is considered the largest but the maximum this species is can measure 70Â cm (28Â in) and a disk diameter of about 14.3Â cm (5.6Â in). Outside from euryalids, the biggest ophiurid brittle star may be Ophiopsammus maculata (6-7 inches).
- Sea stars (Asteroidea)
- The heaviest sea star is Thromidia gigas from the Indo-Pacific, which can surpass 6Â kg (13Â lb) in weight, but only has a diameter of about 65Â cm (2.13Â ft). Despite its relatively small disk and weight, the long slender arms of Midgardia xandaros from the Gulf of California makes it the sea star with the largest diameter at about 1.4Â m (4.5Â ft). Mithrodia clavigera may also become wider than 1Â m (39Â in) in some cases, with stout arms.
Ribbon worms (Nemertea)
The largest nemertean is the bootlace worm, Lineus longissimus. A specimen found washed ashore on a beach in St. Andrews, Scotland in 1864 was recorded at a length of 55Â m (180Â ft).
Mollusks (Mollusca)
Both the largest mollusks and the largest of all invertebrates (in terms of mass) are the largest squids. The colossal squid (Mesonychoteuthis hamiltoni) is projected to be the largest invertebrate. Current estimates put its maximum size at 12 to 14Â m (39 to 46Â ft) long and 750Â kg (1,650Â lb), based on analysis of smaller specimens. In 2007, authorities in New Zealand announced the capture of the largest known colossal squid specimen. It was initially thought to be 10Â m (33Â ft) and 450Â kg (990Â lb). It was later measured at 4.2Â m (14Â ft) long and 495Â kg (1,091Â lb) in weight. The mantle was 2.5Â m (8.2Â ft) long when measured.
The giant squid (Architeuthis dux) was previously thought to be the largest squid, and while it is less massive and has a smaller mantle than the colossal squid, it may exceed the colossal squid in overall length including tentacles. One giant squid specimen that washed ashore in 1878 in Newfoundland reportedly measured 18Â m (59Â ft) in total length (from the tip of the mantle to the end of the long tentacles), head and body length 6.1Â m (20Â ft), 4.6Â m (15Â ft) in diameter at the thickest part of mantle, and weighed about 900Â kg (2,000Â lb). This specimen is still often cited as the largest invertebrate that has ever been examined. However, no animals approaching this size have been scientifically documented and, according to giant squid expert Steve O'Shea, such lengths were likely achieved by greatly stretching the two tentacles like elastic bands.
- Aplacophorans (Aplacophora)
- The largest known of these worm-like, shell-less mollusks are represented in the genus Epimenia, which can reach 30Â cm (12Â in) long. Most aplacophorans are less than 5Â cm (2.0Â in) long.
- Chitons (Polyplacophora)
- The largest of the chitons is the gumboot chiton, Cryptochiton stelleri, which can reach a length of 33Â cm (13Â in) and weigh over 2Â kg (4.4Â lb).
- Bivalves (Bivalvia)
- The largest of the bivalve mollusks is the giant clam, Tridacna gigas. Although even larger sizes have been reported for this passive animal, the top verified size was for a specimen from the Great Barrier Reef. This creature weighed 270Â kg (600Â lb), had an axial length of 1.14Â m (3.7Â ft) and depth of 0.75Â m (2.5Â ft). The largest bivalve ever was Platyceramus platinus, a Cretaceous giant that reached an axial length of up to 3Â m (nearly 10Â ft).
- Gastropods (Gastropoda)
- The "largest" of this most diverse and successful mollusk class of slugs and snails can be defined in various ways.
- The living gastropod species that has the largest (longest) shell is Syrinx aruanus with a maximum shell length of 0.91Â m (3.0Â ft), a weight of 18Â kg (40Â lb) and a width of 96Â cm (38Â in). Another giant species is Melo amphora, which in a 1974 specimen from West Australia, measured 0.71Â m (2.3Â ft) long, had a maximum girth of 0.97Â m (3.2Â ft) and weighed 16Â kg (35Â lb).
- The largest shell-less gastropod is the giant black sea hare (Aplysia vaccaria) at 0.99Â m (3.2Â ft) in length and almost 14Â kg (31Â lb) in weight.
- The largest of the land snails is the giant African snail (Achatina achatina) at up to 1Â kg (2.2Â lb) and 35Â cm (14Â in) long.
- Cephalopods (Cephalopoda)
- (See Cephalopod size.) While generally much smaller than the giant Architeuthis and Mesonychoteuthis, the largest of the octopuses, the giant Pacific octopus (Enteroctopus dofleini), can grow to be very large. The largest confirmed weight of a giant octopus is 74Â kg (163Â lb), with a 7Â m (23Â ft) arm span (with the tentacles fully extended) and a head-to-tentacle-tip length of 3.9Â m (13Â ft). Specimens have been reported up to 125Â kg (276Â lb) but are unverified.
Velvet worms (Onychophora)
Solórzano's velvet worm (Peripatus solorzanoi) is the largest velvet worm known. An adult female was recorded to have a body length of 22 cm (approximately 8.7 in).
Arthropods (Arthropoda)
The largest arthropod known to have existed is the eurypterid (sea scorpion) Jaekelopterus, reaching up to 2.5Â m (8.2Â ft) in body length, followed by the millipede relative Arthropleura at around 2.1Â m (6.9Â ft) in length. Among living arthropods, the Japanese spider crab (Macrocheira kaempferi) is the largest in overall size, the record specimen, caught in 1921, had an extended arm span of 3.8Â m (12Â ft) and weighed about 19Â kg (42Â lb). The heaviest is the American lobster (Homarus americanus), the largest verified specimen, caught in 1977 off of Nova Scotia weighed 20Â kg (44Â lb) and its body length was 1.1Â m (3.6Â ft). The largest land arthropod and the largest land invertebrate is the coconut crab (Birgus latro), up to 40Â cm (1.3Â ft) long and weighing up to 4Â kg (8.8Â lb) on average. Its legs may span 1Â m (3.3Â ft).
Arachnids (Arachnida)
- Both spiders and scorpions include contenders for the largest arachnids.
- Spiders (Araneae)
- The largest species of arachnid by length is probably the giant huntsman spider (Heteropoda maxima) of Laos, which in 2008 replaced the Goliath birdeater (Theraphosa blondi) of northern South America as the largest known spider by leg-span. However the most massive arachnids, of comparable dimensions and possibly even greater mass, are the Chaco golden knee, Grammostola pulchripes, and the Brazilian salmon pink, Lasiodora parahybana. The huntsman spider may span up to 29Â cm (11Â in) across the legs, while in the New World "tarantulas" like Theraphosa can range up to 26Â cm (10Â in). In Grammostola, Theraphosa and Lasiodora, the weight is projected to be up to at least 150Â g (5.3Â oz) and body length is up to 10Â cm (3.9Â in).
- Scorpions (Scorpiones)
- The largest of the scorpions is the species Heterometrus swammerdami of the Indian subcontinent, which have a maximum length of 29.2Â cm (11.5Â in) and weigh around 60Â g (2.1Â oz). Another extremely large scorpion is the African emperor scorpion (Pandinus imperator), which can weigh 57Â g (2.0Â oz) but is not known to exceed a length of 23Â cm (9.1Â in). However, they were dwarfed by Pulmonoscorpius kirktonensis, a giant extinct species of scorpion from Scotland, at an estimated length of 0.7Â m (2.3Â ft) and weight of 15Â kg (33Â lb), and the aquatic Brontoscorpio, at up to 1Â m (3.3Â ft) and a similar weight.
- Pseudoscorpions (Pseudoscorpiones))
- The largest pseudoscorpion is Garypus titanius, from Ascension island, which can be 12Â mm (0.47Â in) long.
Crustaceans (Crustacea)
- The largest crustaceans are crab Tasmanian giant crab (''Pseudocarcinus gigas) 13 kilograms (29Â lb) and a carapace width of up to 46 centimetres (18Â in). It is the only species in the genus Pseudocarcinus. Males reach more than twice the size of females. It has a white shell with claws that are splashed in red. The females' shells change colour when they are producing eggs. Lysiosquillina maculataAt a length of up to 40Â cm, L. maculata is the largest mantis shrimp in the world. L. maculata may be distinguished from its congener L. sulcata by the greater number of teeth on the last segment of its raptorial claw, and by the colouration of the uropodal endopod, the distal half of which is dark in L. maculata but not in L. sulcata. There is a small artisanal fishery for this species. are largest shrimps Tasmanian giant freshwater crayfish (Astacopsis gouldi) 5 kilograms (11Â lb) in weight and over 80 centimetres (31Â in) long have been known in the past, but now, even individuals over 2 kilograms (4.4Â lb) are rare. The species is only found in Tasmanian rivers flowing north into the Bass Strait below 400 metres (1,300Â ft) above sea level, and is listed as an endangered species on the IUCN Red List.
- Branchiopods (Branchiopoda)
- The largest of these primarily freshwater crustaceans is probably Branchinecta gigas, which can reach a length 10Â cm (3.9Â in).
- Barnacles and allies (Maxillopoda)
- The largest species is a copepod (Pennella balaenopterae), known exclusively as a parasite from the backs of fin whales (Balaenoptera physalus). The maximum size attained is 32Â cm (about 13Â in). The largest of the barnacles is the giant acorn barnacle, Balanus nubilis, reaching 7Â cm (2.8Â in) in diameter and 12.7Â cm (5.0Â in) high.
- Ostracods (Ostracoda)
- The largest living representative of these small and little-known but numerous crustaceans is the species Gigantocypris australis females of which reaching a maximum length of 3Â cm (1.2Â in).
- Amphipods, isopods, and allies (Peracarida)
- The largest species is the giant isopod (Bathynomus pergiganteus), which can reach a length of 45Â cm (18Â inches) and a weight of 1.7Â kg (3.7Â lb).
- Remipedes (Remipedia)
- The largest of these cave-dwelling crustaceans is the species Godzillius robustus, at up to 4.5Â cm (1.8Â in).
Horseshoe crabs (Merostomata)
- The four modern horseshoe crabs are of roughly the same sizes, with females measuring up to 60Â cm (2.0Â ft) in length and 5Â kg (11Â lb) in weight.
Sea spiders (Pycnogonida)
- The largest of the sea spiders is the deep-sea species Colossendeis colossea, attaining a leg span of nearly 60Â cm (2.0Â ft).
Trilobites (Trilobita)
- Some of these extinct marine arthropods exceeded 60Â cm (24Â in) in length. A nearly complete specimen of Isotelus rex from Manitoba attained a length over 70Â cm (28Â in), and an Ogyginus forteyi from Portugal was almost as long. Fragments of trilobites suggest even larger record sizes. An isolated pygidium of Hungioides bohemicus implies that the full animal was 90Â cm (35Â in) long.
Myriapods (Myriapoda)
- Centipedes (Chilopoda)
- The biggest of the centipedes is Scolopendra gigantea of the neotropics, reaching a length of 33Â cm (13Â in).
- Millipedes (Diplopoda)
- Two species of millipede both reach a very large size Archispirostreptus gigas of East Africa and Scaphistostreptus seychellarum, endemic to the Seychelles islands. Both of these species can slightly exceed a length of 28Â cm (11Â in) and measure over 2Â cm (0.79Â in) in diameter. The largest ever known was the Arthropleura, a gigantic prehistoric specimen that reached nearly 6.2 feet.
Insects (Insecta)
Insects, a class of Arthropoda, are easily the most numerous class of organisms, with over one million identified species, and probably many undescribed species. The heaviest insect is almost certainly a species of beetle, which incidentally is the most species-rich order of organisms. Although heavyweight giant wetas (Deinacrida heteracantha) are known, the elephant beetles of Central and South America, (Megasoma elephas) and (M. actaeon), the Titan beetle (Titanus giganteus) of the neotropical rainforests or the Goliath beetles, (Goliathus goliatus) and (G. regius), of Africa's rainforests are thought to reach a higher weight. The most frequently crowned are the Goliath beetles, the top known size of which is at least 100Â g (3.5Â oz) and 11.5Â cm (4.5Â in). The elephant beetles and titan beetle can reach greater lengths than the Goliath, at up to 13.1 and 15.2Â cm (5.2 and 6.0Â in), respectively, but this is in part thanks to their rather large horns. The Goliath beetle's wingspan can range up to 25Â cm (9.8Â in).
Some moths and butterflies have much larger areas than the heaviest beetles, but weigh a fraction as much.
The longest insects are the stick insects, see below.
Representatives of the extinct dragonfly-like order Protodonata such as the Carboniferous Meganeura monyi of what is now France and the Permian Meganeuropsis permiana of what is now North America are the largest insect species yet known to have existed. These creatures had a wingspan of some 75Â cm (30Â in) and a mass of over 1 pound (450Â g), making them about the size of a crow.
- Cockroaches and termites (Blattodea)
- The largest cockroach is the Australian giant burrowing cockroach (Macropanesthia rhinoceros). This species can attain a length of 8.3Â cm (3.3Â in) and a weight of 36Â g (1.3Â oz). The giant cockroach (Blaberus giganteus) of the neotropics reaches comparable lengths although is not as massive as the burrowing species. The termites, traditionally classified in their own order (Isoptera), have recently been re-considered to belong in Blattodea. The largest of the termites is the African species Macrotermes bellicosus. The queen of this species can attain a length of 14Â cm (5.5Â in) and breadth of 5.5Â cm (2.2Â in) across the abdomen; other adults, on the other hand, are about a third of the size.
- Beetles (Coleoptera)
- The beetles are the largest order of organisms on earth, with about 400,000 species so far identified. The most massive species are the Goliathus, Megasoma and Titanus beetles already mentioned. Another fairly large species is the Hercules beetle (Dynastes hercules) of the neotropic rainforests with a maximum overall length of at least 19Â cm (7.5Â in) including the extremely long pronotal horn. The weight in this species does not exceed 16.5Â g (0.58Â oz). The longest overall beetle is a species of longhorn beetle, Batocera wallacei, from New Guinea, which can attain a length of 26.6Â cm (10.5Â in), about 19Â cm (7.5Â in) of which is comprised by the long antennae.
- Earwigs (Dermaptera)
- The largest of the earwigs is the Saint Helena giant earwig (Labidura herculeana), endemic to the island of its name, which is up to 8Â cm (3.1Â in) in length.
- True flies (Diptera)
- The largest species of this order, which includes the common housefly, is the neotropical species Gauromydas heros, which can reach a length of 6Â cm (2Â 3â8Â in) and a wingspan of 10Â cm (3.9Â in). Species of crane fly, the largest of which is Holorusia brobdignagius, can attain a length of 23Â cm (9.1Â in) but are extremely slender and much lighter in weight than Gauromydas.
- Mayflies (Ephemeroptera)
- The largest mayflies are members of the genus Proboscidoplocia from Madagascar. These insects can reach a length of 7Â cm (2.8Â in).
- True bugs (Hemiptera)
- The largest species of this diverse order is usually listed as the giant water bug in the genus Lethocerus, with L. maximus from the Neotropics being the absolutely largest. They can surpass 12Â cm (4.7Â in) in length, with some suggesting that the maximum size is 15Â cm (5.9Â in). It is more slender and less heavy than most other insects of this size (principally the huge beetles). The largest cicada is Megapomponia imperatoria, which has a head-body length of about 7Â cm (2.8Â in) and a wingspan of 18â"20Â cm (7â"8Â in). The cicadas of the genus Tacua can also grow to comparably large sizes. The largest type of aphid is the giant oak aphid (Stomaphis quercus), which can reach an overall length of 2Â cm (0.79Â in). The biggest species of leafhopper is Ledromorpha planirostris, which can reach a length of 2.8Â cm (1.1Â in).
- Ants and allies (Hymenoptera)
- The largest of the ants, and the heaviest species of the order, are the females of the African Dorylus helvolus, reaching a length of 5.1Â cm (2.0Â in) and a weight of 8.5Â g (0.30Â oz). The ant that averages the largest for the mean size within the whole colony is a ponerine ant, Dinoponera gigantea, from South America, averaging up to 3.3Â cm (1.3Â in) from the mandibles to the end of abdomen. Workers of the bulldog ant (Myrmecia brevinoda) of Australia are up to 3.7Â cm (1.5Â in) in total length, although much of this is from their extremely large mandibles. The largest of the bee species, also in the order Hymenoptera, is Megachile pluto of Indonesia, the females of which can be 3.8Â cm (1.5Â in) long, with a 6.3Â cm (2.5Â in) wingspan. Nearly as large, the carpenter bees can range up to 2.53Â cm (1.00Â in). The largest wasp is probably the so-called tarantula hawk species Pepsis pulszkyi of South America, at up to 6.8Â cm (2.7Â in) long and 11.6Â cm (4.6Â in) wingspan, although many other Pepsis approach a similar size.
- Moths and allies (Lepidoptera)
- The Hercules moth (Coscinocera hercules), in the family Saturniidae, is endemic to New Guinea and northern Australia, and its wings have the largest documented surface area (300 square centimeters) of any living insect, and a maximum wingspan which is confirmed to 28Â cm (11Â in) while unconfirmed specimens have spanned up to 35.5Â cm (14.0Â in). The largest species overall is often claimed to be either the Queen Alexandra's birdwing (Ornithoptera alexandrae), a butterfly from Papua New Guinea, or the Atlas moth (Attacus atlas), a moth from Southeast Asia. Both of these species can reach a length of 8Â cm (3.1Â in), a wingspan of 28Â cm (11Â in) and a weight of 12Â g (0.42Â oz). One Atlas moth allegedly had a wingspan of 30Â cm (12Â in) but this measurement was not verified. The larvae in the previous species can weigh up to 58 and 54Â g (2.0 and 1.9Â oz), respectively. However, there are no reported measurements of surface area that would exceed the Hercules moth, and the white witch (Thysania agrippina) of Central and South America, has the largest recorded wingspan of the order, and indeed of any living insect, though the white witch is exceeded in surface area by the Hercules moth. The verified record-sized Thysania spanned 30.8Â cm (12.1Â in) across the wings, although specimens have been reported to 36Â cm (14Â in). The heaviest mature moths have been cited in the giant carpenter moth (Xyleutes boisduvali) of Australia, which has weighed up to 20Â g (0.71Â oz) although the species does not surpass 25.5Â cm (10.0Â in) in wingspan.
- Mantises (Mantodea)
- The largest species of this order is Toxodera denticulata from Java, which has been measured up to 20Â cm (7.9Â in) in overall length. However, an undescribed species from the Cameroon jungle is allegedly much larger than any other mantis and may rival the larger stick insects for the longest living insect. Among widespread mantis species, the largest is the Chinese mantis (Tenodera aridifolia). The females of this species can attain a length of up to 10.6Â cm (4.2Â in).
- Alderflies and allies (Megaloptera)
- This relatively small insect order includes some rather large species, many of which are noticeable for their elongated, imposing mandibles. The dobsonflies reach the greatest sizes of the order and can range up to 12.5Â cm (4.9Â in) in length.
- Net-winged insects (Neuroptera)
- These flying insects reach their largest size in Palparellus voeltzkowi, which can have a wingspan over 16Â cm (6.3Â in). The largest lacewing is the "blue eyes lacewing" (Nymphes myrmeleonides) of Australia, which can measure up to 4Â cm (1.6Â in) in length and span 11Â cm (4.3Â in) across the wings. Some forms of this ancient order could grow extremely large during the Jurassic Era and may have ranked among the largest insects ever.
- Dragonflies (Odonata)
- The largest living species of dragonfly is Megaloprepus caerulatus of the neotropics, attaining a size of as much as 19Â cm (7.5Â in) across the wings and a body length of over 12Â cm (4.7Â in). Spanning up to 17.6Â cm (6.9Â in) and measuring up to 11.8Â cm (4.6Â in) long, Tetracanthagyna plagiata of Southeast Asia is bulkier and heavier than Megaloprepus at up to 7Â g (0.25Â oz). The largest species of dragonfly ever is the extinct aforementioned Meganeura, although it is not certain to be included in the modern dragonfly order.
- Grasshoppers and allies (Orthoptera)
- The largest of this widespread, varied complex of insects are the giant wetas of New Zealand, which is now split among 12 species. The largest of these is the Little Barrier Island giant weta (Deinacrida heteracantha), the largest specimen was weighed at 71.3Â g (2.52Â oz), one of the largest insects weights ever known. These heavyweight insects can be over 9Â cm (3.5Â in) long. The largest grasshopper species is often considered to be the Australian giant grasshopper (Valanga irregularis), which ranges up to 9Â cm (3.5Â in) in length. The American eastern lubber grasshopper (Romalea guttata) can allegedly range up to 10Â cm (3.9Â in) in length. However, the greatest grasshopper sizes known, to 12Â cm (4.7Â in), have been cited in the South American giant grasshopper (Tropidacris violaceus). The longest members of this order (although much lighter than the giant wetas) is the katydid Macrolyristes corporalis of Southeast Asia which can range up to 21.5Â cm (8.5Â in) with its long legs extended and can have a wingspan of 20Â cm (7.9Â in).
- Stick insects (Phasmatodea)
- The longest known stick insect, and indeed the longest insect ever known, is Phobaeticus chani of the Bornean rainforests, with one specimen held in the Natural History Museum in London measuring 56.7Â cm (22.3Â in) in total length. This measurement is, however, with the front legs fully extended. The body alone still measures 35.7Â cm (14.1Â in). The species with the second longest body is Phobaeticus kirbyi, also of Borneo, which measures up to 32.8Â cm (12.9Â in), while the overall length (from the hind to the front legs) is up to 54.6Â cm (21.5Â in). The second longest insect in terms of total length is Phobaeticus serratipes of Malaysia and Singapore, measuring up to 55.5Â cm (21.9Â in). Another extremely long stick insect is Pharnacia maxima, which measured 51Â cm (20Â in) with its legs extended. The spiny stick insect (Heteropteryx dilatata) of Malaysia does not reach the extreme lengths of its cousins, the body reaching up to 16Â cm (6.3Â in) long, but it is much bulkier. The largest Heteropteryx weighed about 65Â g (2.3Â oz) and was 3.5Â cm (1.4Â in) wide across the thickest part of the body.
- Lice (Phthiraptera)
- These insects, which live parasitically on other animals, are as a rule quite small. The largest known species is the hog louse, Haematopinus suis, a sucking louse that lives on large livestock like pigs and cattle. It can range up to 6Â mm (0.24Â in) in length.
- Stoneflies (Plecoptera)
- The largest species of stonefly is Pteronarcys californica of western North America, a species favored by fishermen as lures. This species can attain a length of 5Â cm (2.0Â in) and a wingspan of over 9.5Â cm (3.7Â in).
- Booklice (Psocoptera)
- The largest of this order of very small insects are the barklice of the genus Psocus, the top size of which is about 1 cm.
- Fleas (Siphonaptera)
- The largest species of flea is Hystrichopsylla schefferi. This parasite is known exclusively from the fur of the mountain beaver (Aplodontia rufa) and can reach a length of 1.2Â cm (0.47Â in).
- Silverfishes and allies (Thysanura)
- These strange-looking insects, known to feed on human household objects, can range up to 4.3Â cm (1.7Â in) in length. A 350 million year old form was known to grow quite large, at up to 6Â cm (2.4Â in).
- Thrips (Thysanoptera)
- Members of the genus Phasmothrips are the largest kinds of thrips. The maximum size these species attain is approximately 1.3Â cm (0.51Â in) in length.
- Caddisflies (Trichoptera)
- The largest of the small, moth-like caddisflies is Eubasilissa maclachlani. This species can range up to 7Â cm (2.8Â in) across the wings.
Fungi
The largest living fungus may be a honey fungus of the species Armillaria ostoyae. A mushroom of this type in the Malheur National Forest in the Blue Mountains of eastern Oregon, U.S. was found to be the largest fungal colony in the world, spanning 8.9Â km2 (2,200 acres) of area. This organism is estimated to be 2,400 years old. The fungus was written about in the April 2003 issue of the Canadian Journal of Forest Research. While an accurate estimate has not been made, the total weight of the colony may be as much as 605 tons. If this colony is considered a single organism, then it is the largest known organism in the world by area, and rivals the aspen grove "Pando" as the known organism with the highest living biomass. It is not known, however, whether it is a single organism with all parts of the mycelium connected.
In Armillaria ostoyae, each individual mushroom (the fruiting body, similar to a flower on a plant) has only a 5Â cm (2.0Â in) stipe, and a pileus up to 12.5Â cm (4.9Â in) across. There are many other fungi which produce a larger individual size mushroom. The largest known fruiting body of a fungus is a specimen of Phellinus ellipsoideus (formerly Fomitiporia ellipsoidea) found on Hainan Island. The fruiting body masses up to 500Â kg (1,100Â lb).
Until P. ellipsoideus replaced it, the largest individual fruit body came from Rigidoporus ulmarius. R. ulmarius can grow up to 284Â kg (626Â lb), 1.66Â m (5.4Â ft) tall, 1.46Â m (4.8Â ft) across, and has a circumference of up to 4.9Â m (16Â ft).
Protists
(Note: the group Protista is not used in current taxonomy.)
Amoebozoans (Amoebozoa)
- Among the organisms that are not multicellular, the largest are the slime moulds, such as Physarum polycephalum, some of which can reach a diameter over 30Â cm (12Â in). These organisms are unicellular, but they are multinucleate.
Euglenozoans (Euglenozoa)
- Some euglenophytes, such as certain species of Euglena, reach lengths of 400 μm.
Rhizarians (Rhizaria)
- The largest species traditionally considered protozoa are giant amoeboids like foraminiferans. One such species, the xenophyophore Syringammina fragilissima, can attain a size of 20Â cm (7.9Â in).
Alveolates (Alveolata)
- The largest ciliates, such as Spirostomum, can attain a length over 4Â mm (0.16Â in).
Stramenopiles (Stramenopila)
- The largest stramenopiles are giant kelp from the northwestern Pacific. The floating stem of Macrocystis pyrifera can grow to a height of over 45Â m (148Â ft).
- Macrocystis also qualifies as the largest brown alga, the largest chromist, and the largest protist generally.
Bacteria
The largest known species of bacterium is Thiomargarita namibiensis, which grows to 0.75Â mm (0.030Â in) in diameter, making it visible to the naked eye and a thousand times the size of more typical bacteria.
- Cyanobacteria
- One of the largest "blue green algae" is Lyngbya, whose filamentous cells can be 50 μm wide.
Viruses
The largest virus on record so far is the Pithovirus sibericum with the length of 1.5 micrometres, comparable to the typical size of a bacterium and large enough to be seen in light microscopes. It was discovered in March 2014 in a soil sample collected from a riverbank in Siberia. Prior to this discovery, the largest virus was the peculiar Pandoravirus genus of viruses, which have a size of approximately 1 micrometer and whose genome contains 1,900,000 to 2,500,000 base pairs of DNA.
Both these viruses infect amoebas specifically.
See also
References
External links
- 10 of the largest living things on the planet Melissa Breyer. TreeHugger April 28, 2015